Polymorphism may play a role in speciation, but how morphs diversify into independent lineages is unclear. The alternative adaptation hypothesis (West-Eberhard 1986) predicts that 'developmental character release', generated by the loss of a morph, plays a key role.

We are investigating this hypothesis by comparing body size, and wing shape in closely related damselflies that co-exist as either polymorphic or monomorphic populations. This work is part of a long-term collaboration with Prof. Yoshitaka Tsubaki (Kyoto).

In a more recent collaboration with Dr Phill Watts (Oulu, Finland) we are hoping to carry out out a phylogenetic appraisal of Mnais damselflies throughout Eastern Asia, with emphasis on Japanese populations.